All land tortoises are herbivorous in the wild. The first turtles already existed in the era of the dinosaurs, some 300 million years ago. Turtles and tortoises are the only surviving branch of the even more ancient clade Anapsida, which includes groups such as the procolophonoids, millerettids and pareiasaurs. Most of the anapsids became extinct in the late Permian period, with the exception of the procolophonoids and the precursors of the testudines (turtles and tortoises).














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